Encephalartos cupidus

Assessor: Sarah Schumann

Sensitive in 2010
No
Family
Zamiaceae
Reason for the sensitivity status
This taxon is sensitive as it is threatened with illegal collection of wild individuals, causing population decline whereby it is considered extinct in most of its geographic range. Evidence shows that the Encephalartos genus is in high demand in international trade for ornamental and horticultural purposes as well as local medicinal purposes. The species has a small remaining population size making it vulnerable to further population loss. Releasing data on this species can exacerbate threat and vulnerability.
This species is extremely rare in the wild and is known to be exploited, utilised or traded. The localities of remaining populations need to be protected to avoid any further exploitation, which is likely to drive it to extinction.
Exploitation extent
Significant - wild individuals of the species are known to be exploited, collected, traded or utilized in a targeted manner, and utilisation is widespread, affects the majority of wild populations and/or is causing rapid decline of the wild population.
Justification and references

All South African cycads as well as this species is included in Appendix I of the CITES Convention which lists all species threatened with extinction that are, or may be affected by trade. According to the SANBI Red List Assessment states, this species is Critically Endangered as it is majorly threatened by over-collecting for ornamental purposes and medicinal use resulting in reproductive failure (Donaldson 2010; Donaldson 2003). It is extinct throughout most of its original geographic extent with only a few individuals remaining in the wild and has declined by 80% in the past 50 years (Donaldson 2010). Population decline is high and habitat destruction is moderate (Donaldson 2003). Cycad species are in trade demand and at risk from illegal collection (Smith 2014; Yeld 2014). Studies present strong evidence of ongoing and accelerating rates of decline in most cycad species due mainly to illegal removal and this is especially evident with endangered and critically endangered Encephalartos species (Okubamichael et al. 2016; Cousins & Witkowski 2017). Research by Cousins (2012) shows that this species is among the 25 of 37 Encephalartos species that is being utilised for traditional medicine. The CITES Trade Database recorded the trade of the species for commercial, personal, scientific purposes. Sourced from artificial propagation between 1986-2019. 

CITES trade statistics derived from the CITES Trade Database, UNEP World Conservation Monitoring Centre, Cambridge, UK.

Cousins, S.R., Williams, V.L., & Witkowski, E.T.F. 2012. Uncovering the cycad taxa (Encephalartos species) traded for traditional medicine in Johannesburg and Durban, South Africa. South African Journal of Botany78, 129–138. DOI:10.1016/j.sajb.2011.06.001

Cousins, S.R. and Witkowski, E.T.F., 2017. African cycad ecology, ethnobotany and conservation: a synthesis. The Botanical Review83(2), pp.152-194.

Donaldson, J.S. 2003. Cycads. Status survey and conservation action plan. IUCN/SSC Cycad Specialist Group, Gland, Switzerland; Cambridge, UK.

Donaldson, J.S. 2010. Encephalartos cupidus R.A.Dyer. National Assessment: Red List of South African Plants version 2020.1. 

Okubamichael, D. Y., S. L. Jack, J. D. Bösenberg, M. T. Hoffman & J. S. Donaldson. 2016. Repeat photography confirms alarming decline in south African cycads. Biodiversity Conservation 25(11): 2153–2170.

Smith, D. 2014. 1 September 2014. South Africa's ancient cycad plants under threat from poachers. The Guardian. Available at: https://www.theguardian.com/world/2014/sep/01/ancient-cycad-plants-under-threat-poachers

Yeld, J. 2014. 7 August 2014. Cycad thieves strike again. IOL. Available at: https://www.iol.co.za/news/cycad-thieves-strike-again-1731799

Population vulnerability
Population is vulnerable: size is <= 2500 mature individuals OR the number of known subpopulations is <= 5 OR range is <= 100km2 OR species at risk of localised extinctions
Justification and references

The known extent occurrence is 20 km² and area of occupancy is 10 km² (Donaldson 2010). Currently there are 50 plants remaining in the wild (Bösenberg. DW, personal communication 2021, 1 December)

Donaldson, J.S. 2010. Encephalartos cupidus R.A.Dyer. National Assessment: Red List of South African Plants version 2020.1. 

Donaldson, J.S. 2010. Encephalartos cupidusThe IUCN Red List of Threatened Species 2010: e.T41884A10567684. https://dx.doi.org/10.2305/IUCN.UK.2010-3.RLTS.T41884A10567684.en.

Targeted demographics
Mature (breeding) individuals are killed, significantly weakened or are permanently removed from the wild, OR immature individuals are targeted and this significantly impacts mature (breeding) individuals.
Justification and references

According to the CITES Trade Database, mature individuals, seeds and leaves are known to be collected and traded. Threat of illegal removal of individuals have caused population decline (Donaldson 2010; Donaldson 2003).

CITES trade statistics derived from the CITES Trade Database, UNEP World Conservation Monitoring Centre, Cambridge, UK.

Donaldson, J.S. 2003. Cycads. Status survey and conservation action plan. IUCN/SSC Cycad Specialist Group, Gland, Switzerland; Cambridge, UK.

Donaldson, J.S. 2010. Encephalartos cupidus R.A.Dyer. National Assessment: Red List of South African Plants version 2020.1. 

Regeneration potential
This species has a slow population growth rate, or the growth rate varies depending on habitat, and there is a poor chance the wild populations will recover from exploitation OR a collector might feasibly harvest the entire extant population removing the chance of subsequent recruitment.
Justification and references

Research shows that cycads are generally long-lived, have slow growth rates, produce low numbers of offspring and have infrequent recruitment (Golding & Hurter 2003, Donaldson 2003, Cousins & Witkowski 2017). All groups of Encephalartos experience population decline from removal of adult plants (Donaldson 1995) .Viable seed production in Encephalartos populations tends to decline with decreasing population size (Cousins & Witkowksi 2017). Due to the ongoing over-collection, low number of individuals and restricted range this species has lost majority of its population (Donaldson 2010). The remaining population of this species occurs in a private nature reserve (Bösenberg. DW, personal communication 2021, 1 December), making it highly vulnerable to further population loss were exploitation to occur on this site too. Recruitment and recovery may be poor. 

Cousins, S.R. and Witkowski, E.T.F., 2017. African cycad ecology, ethnobotany and conservation: a synthesis. The Botanical Review83(2), pp.152-194.

Donaldson, J.S., 1995. Understanding cycad life histories, an essential basis for successful conservation. In: Donaldson, J.S. (Ed.), Cycad Conservation in South Africa, Issues, Priorities and Actions. The Cycad Society of South Africa, Stellenbosch, pp. 8–13.

Donaldson, J.S. 2003. Cycads. Status survey and conservation action plan. IUCN/SSC Cycad Specialist Group, Gland, Switzerland; Cambridge, UK.

Donaldson, J.S. 2010. Encephalartos cupidus R.A.Dyer. National Assessment: Red List of South African Plants version 2020.1. 

Golding, J.S. and Hurter, P.J.H., 2003. A Red List account of Africa's cycads and implications of considering life-history and threats. Biodiversity & Conservation12(3), pp.507-528.